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Akçay, E., & Van Cleve, J. (2012). Behavioral responses in structured populations pave the way to group optimality. The American Naturalist, 179(2), 257-69. https://doi.org/10.1086/663691

Akçay, E., Van Cleve, J., Feldman, M., & Roughgarden, J. (2009). A theory for the evolution of other-regard integrating proximate and ultimate perspectives. Proceedings of the National Academy of Sciences of the United States of America, 106(45), 19061-6. https://doi.org/10.1073/pnas.0904357106 (Original work published 2009)

Akçay, E., & Van Cleve, J. (2016). There is no fitness but fitness, and the lineage is its bearer. Philosophical Transactions of the Royal Society of London. Series B, Biological Sciences, 371(1687), 20150085. https://doi.org/10.1098/rstb.2015.0085 (Original work published 2016)

Lehmann, L., Mullon, C., Akçay, E., & Van Cleve, J. (2016). Invasion fitness, inclusive fitness, and reproductive numbers in heterogeneous populations. Evolution; International Journal of Organic Evolution, 70(8), 1689-702. https://doi.org/10.1111/evo.12980

McGlothlin, J., Akçay, E., Brodie, E., Moore, A., & Van Cleve, J. (2022). A Synthesis of Game Theory and Quantitative Genetic Models of Social Evolution. The Journal of Heredity, 113(1), 109-119. https://doi.org/10.1093/jhered/esab064 (Original work published 2022)